Mitsuha Miyamizu, a high school girl, wishes to live the life of a male in Tokyo, a fantasy that contrasts sharply with her current existence in the countryside. Meanwhile, Taki Tachibana, a high school student balancing a part-time job and dreams of a career in design, lives a hectic existence in the city. Mitsuha awakens one day in a room that isn't her own and finds herself enjoying the dream life in Tokyo—but in Taki's body! Taki, on the other hand, finds himself living Mitsuha's life in the impoverished countryside. In quest of an explanation for this bizarre occurrence, they begin to look for one another.